In the era known as "New A.D., six months after the "DC War" against the military organization "Divine Crusaders', and the "L5 Campaign" against the extraterrestrial foes known as "Aerogaters"... Lt. Kyosuke Nanbu's Earth Federation PT Squad "The ATX Team" heads to the far east to defeat the survivors of DC. In the middle of the fight, he and his partner Excellen witness the sudden arrival of a new humanoid weapon. This new angel-like machine, is it a friend or a foe

In the middle of the battle with the remnants of DC, a mysterious woman named Lamia Loveless appears, piloting a Special Unit (A Special Humanoid Mobile Weapon) She has strange problems with her language, and her unknown origins make Kyosuke suspicious. Meanwhile, Archibold's DC forces attack a dig site in China called Shiyu Mound. The federation battlecarrier Shirogane rushes to intercept, and the ATX team, now with Lamia, sorties. Excellent and Bullet are surprised by the strange feeling in the air around the mound. Then strange plantlike monsters appear, as well as a Tiger and Dragon Chokijin

A mock battle betweent two prototypes is underway at Izu base. Ryusei of the SRX team is piloting a simplified mass produced version of his own R-1, the R-Blade, and Latouni of the special tactics instruction squad is piloting the Wild Falken, the new unit from the ATX plan. Meanwhile, Archibold's DC forces send Yuki's squad to recover the prototype. Two of the members of his squad are Arado and Seolla, young pilots trained at a special institute called "The School". Latooni, who once attended the school herself, encounters Seolla on the battlefield.

To Latooni, Arado and Seolla are old friends fom the Special training organization "The School". Ryusei and the others meet up with the ATX team, and persue the DC remnants. They're stopped by Seolla in the Wild Falken, and Latooni goes after her, but..

The Shirogane is damaged in its fight with the DC remnants. Kyosuke begins to wonder at the enemy's overwhelming firepower, and suspect the existance of a 3rd party behind the scenes. Meanwhile, at the massive underground Earth Cradle, DC's headquarters, forces are steadily gathering..