Claude's dream comes true when she moves up from temp to a permanent position as the assistant to news anchor Will Butler. But all is not quite as she had imagined when she realizes that new co-workers Kipp and Lydia will stop at nothing to sabotage her. Meanwhile Claude's trusty friends Ramona and Owen begrudgingly help her transition from the comfort and safety of the fourth floor to the battlefield that is the 22nd floor, while Claude tries to hide her crush on Will Butler, who is just less than perfect.

When Claude is unexpectedly asked out for dinner with Will, she quickly cancels her double-date plan with Ramona. But when the ""date"" turns out to be less of one than she anticipated, Claude finds solace in the company of Lydia -- who assumes that Claude slept with him. Meanwhile, Owen tries to assert authority over Kipp and the two feud over an expensive pen.

Claude sees herself struggling with her moral values when she is forced to lie for Will. She feels bad about covering up one of Will's escapades and lying to Ramona and his wife about it, but it's all good after Claude talks to Ramona. Meanwhile, the employees attend a companywide meeting where they have to pick a "floor buddy", someone from another floor for them to bond with. Lydia is at first annoyed by Owen but later misses him after she realizes he treated her like no one else did. And Ramona and Kipp pretend to dislike each other after having a great time in the Hamptons.

Feeling jealous over Claude being showered by gifts for being Will Butler's assistant, Kipp and Lydia try to break Claude, Ramona and Owen's friendship. Kipp tells Claude that Will doesn't like her having her 4th floor friends around his office, which Claude believes. She politely asks them to stay on their floor, which they take offensively. Now Claude feels bad for sending her friends away, and tries to work her way back into their friendship circle. Meanwhile, Lydia struggles with her (lack of) good sense after she feels guilty for switching Ramona's original Prada hangbag for a cheap imitation. She was trying o save the purse from being ""killed"" by Ramona's exquisite fashion taste and personal alterations on the wonderful Italian leather.

Ramona tells Claude she has to use her vacation days in the next few days or else she'll lose them. Claude refuses to leave Will even for a week, and Ramona (trying to be a good friend and give Claude a break from her overbearing work) sends a memo to Will about Claude's vacation. Forced to take a break, Claude pretends to go to Niagra Falls and sneaks around the office when she meets her temporary replacement – who she thinks might take her job. Ultimately, Will reassures her she doesn't have to worry about her job; he wouldn't trade her for the world. As for Kipp's dream job of becoming an associate producer, well, that will have to wait a little longer. Torturing Claude with lies about Luke's resume might have given Luke a promotion he never expected in such short notice.